Analysis of the topography associated with large vertical strains measured in the upper crust in the Basin and Range and elsewhere in the US Cordillera shows that it varies systematically with upper crustal strain such that it decreases where tectonism has resulted in relative thinning. The magnitude of topographic deflection suggests that the amount of thickening or thinning of the upper crust cannot also be reflected in vertical motions of the Moho discontinuity, nor can the vertical strain be isostatically accommodated by inflow or outflow of asthenosphere, mantle lithosphere or mafic lower crust. Simple isostatic calculations show that the compensating medium had a density probably within 100-200 kg/m of average upper continental crust, thus indicating that the compensating medium lies within the crust and is probably in large part quartzose. This conclusion is in accord with laboratory experiments on the strengths of rocks, and with reflection seismograms in deformed regions which suggest that the Moho is generally flat beneath regions of extreme gradients in upper crustal vertical strain._x000D_
The upper crust floats on a quartzose layer in the mid-crust which, under orogenic conditions, is apparently so weak that it behaves as a relatively inviscid fluid at geologic timescales (> 10,000 a) and subcontinental lengthscales (100-1000 km).